LACM dino camp 3 - Mamenchisaurus and Triceratops 1

Last night London and I spent the night in the Natural History Museum of Los Angeles County (LACM), as part of the Camp Dino overnight adventure. So we got lots of time to roam the exhibit halls when they were–very atypically–almost empty. Above are the museum’s mounted Triceratops–or one of them, anyway–and mounted cast of the Mamenchisaurus hochuanensis holotype, presented in glorious not-stygian-darkness (if you went through the old dino hall, pre-renovation, you know what I mean).

LACM dino camp 1 - dueling dinos

We got there early and had time to roam around the museum grounds in Exposition Park. The darned-near-life-size bronze dinos out front are a minor LA landmark.

LACM dino camp 2 - fountain

The rose garden was already closed, but we walked by anyway, and caught this rainbow in the big fountain.

LACM dino camp 4  - Mamenchisaurus and Triceratops 2After we checked in we had a little time to roam the museum on our own. I’ve been meaning to blog about how much I love the renovated dinosaur halls. The bases are cleverly designed to prohibit people touching the skeletons without putting railings or more than minimal glass in the way, and you can walk all the way around the mounted skeletons and look down on them from the mezzanine–none of that People’s Gloriously Efficient Cattle Chute of Compulsory Dinosaur Appreciation business. Signage is discreet and informative, and so are the handful of interactive gizmos. London and I spent a few minutes using a big touch-screen with a slider that controlled continental drift from the Triassic to the present–a nice example of using technology to add value to an exhibit without taking away from the real stuff that’s on display. There are even a few places to sit and just take it all in. That’s pretty much everything I want in a dinosaur hall.

Also, check out the jumbotron on the left in the above photo. It was running a (blessedly) narration-free video on how fossils are found, collected, prepared, mounted, and studied, on about a five-minute loop. Lots of pretty pictures. Including this next one.

LACM dino camp 5 - big ilium photo

There are a couple of levels of perspective distortion going on here, both in the original photo and in my photo of that photo projected on the jumbotron. Still, I feel confident positing that that is one goldurned big ilium. I’m not going to claim it’s the biggest bone I’ve ever seen–that rarely ends well–but sheesh, it’s gotta be pretty freakin’ big. And apparently a brachiosaurid, or close to it. Never mind, it’s almost certainly an upside-down Triceratops skull. Thanks to Adam Yates for the catch. I will now diminish, and go into the West.

LACM dino camp 6 - ceratopsian skulls

Triceratops, Styracosaurus, and Einiosaurus–collect the whole set!

LACM dino camp 7 - tyrants

Of course, the centerpiece of the second dinosaur hall–and how great is it that there are two!?–is the T. rex trio: baby, juvenile (out of frame to the right), and subadult. Yes, subadult: the “big” one is not as big as the really big rexes, and from the second floor you can see unfused neural arches in some of the caudal vertebrae (many thanks to Ashley Fragomeni for pointing those out to me on a previous visit).

LACM dino camp 8 - baby rex

Awwwww! C’mere, little fella!

LACM dino camp 9 - pneumatic diplodocid caudals

Still, this ain’t Vulgar Overstudied Theropod Picture of the Week. Here are some sweet pneumatic diplodocid caudals in the big wall o’ fossils (visible behind Mamenchisaurus in the overhead photo above). The greenish color is legit–in the Dino Lab on the second floor, they’re prepping a bunch of sauropod elements that look like they were carved out of jade.

Sculpey allosaur claws

Sudden violent topic shift, the reason for which will be become clear shortly: London and I have been sculpting weapons of mass predation in our spare time. In some of the photos you may be able to see his necklace, which has a shark tooth he sculpted himself. Here are a couple of allosaur claws I made–more on those another time.

LACM dino camp 10  - molding and casting

The point is, enthusiasm for DIY fossils is running very high at Casa Wedel, so London’s favorite activity of the evening was molding and casting. Everyone got to make a press mold using a small theropod tooth, a trilobite, or a Velociraptor claw. Most of the kids I overheard opted for the tooth, but London went straight for the claw.

LACM dino camp 11 - raptor claw mold

Ready for plaster! Everyone got to pick up their cast at breakfast this morning, with instructions to let them cure until this evening. All went well, so I’ll spare you a photo of this same shape in reverse.

LACM dino camp 12 - Camp Wedel in the African bush

We were split into three tribes of maybe 30-40 people each, and each tribe bedded down in a different hall. The T. rex and Raptor tribes got the North American wildlife halls, but our Triceratops tribe got the African wildlife hall, which as a place to sleep is about 900 times cooler. Someone had already claimed the lions when we got there, so London picked hyenas as our totem animals.

LACM dino camp 13 - London with ammonite

Lights out was at 10:30 PM, and the lights came back on at 7:00 this morning. Breakfast was out from 7:15 to 8:00, and then we had the museum to ourselves until the public came in at 9:30. So I got a lot of uncluttered photos of stuff I don’t usually get to photograph, like this ammonite. Everyone should have one of these.

LACM dino camp 14 - Wedel boys with Carnotaurus

London’s favorite dino in the museum is Carnotaurus. It’s sufficiently weird that I can respect that choice.

LACM dino camp 15 - London with rexes

Not that there’s anything wrong with the old standards, especially when they’re presented as cleanly and innovatively as they are here.

LACM dino camp 16 - Matt with Argentinosaurus

Finally, the LACM has a no tripod policy, and if they see you trying to carry one in they will make you take it back to your car. At least during normal business hours. But no one searched my backpack when we went in last night, and I put that sucker to some good use. Including getting my first non-bigfoot picture of the cast Argentinosaurus dorsal. It was a little deja-vu-ey after just spending so much time with the giant Oklahoma Apatosaurus–elements of the two animals really are very comparable in size.

If you’re in the LA area and interested in spending a night at the museum–or at the tar pits!–check out the “Overnight Adventures” page on the museum’s website. Cost is $50 per person for members or $55 for non-members, and worth every penny IMHO. It’s one of those things I wish we’d done years ago.

So, this happened today

March 28, 2013

Big Bend Alamosaurus cervical J with Matt for scale

As I noted in a comment on the previous post, titanosaurs have stupid cervicals.

As evidence, here is as gallery of titanosaur cervicals featured previously on SV-POW!.

1. From Whassup with your segmented lamina, Uberabatitan ribeiroi?, an anterior cervical of that very animal, from Salgado and Carvalho (2008: fig. 5). As well as the titular segmented lamina, note the ridiculous ventral positioning of the cervical rib. It’s like it’s trying to be Apatosaurus, but it just doesn’t have the chops.

uberabatitan-cervicals-copy

2. From Mystery of the missing Malawisaurus vertebra, this alleged vertebra of that taxon from Jacobs et al. (1993:fig. 1), which completely fails to resemble all the other cervicals subsequently described from Malawisaurus (see the earlier post for details). Note the crazy sail-like neural spine and super-fat parapophyseal stump.

malawisaurus-1993

3. From Futalognkosaurus was one big-ass sauropod, this completely insane posterior cervical vertebra of Futalognkosaurus in right anterolateral view, with Juan Porfiri (175 cm) for scale. It’s super-tall — much taller than it is wide, and seemingly taller than it is long.

Posterior cervical vertebra of Futalognkosaurus in right anterolateral view; Juan Porfiri (175 cm) for scale

4. From Ch-ch-ch-changes, cervical 11 of Rapetosaurus, from Curry Rogers (2009:fig. 5). Notice how tiny the centrum is compared with the tall superstructure, and how the neural spine has such a distinct peak. Weird.

Rapetosaurus cervical

5. From Talking about sauropods on The Twenty-First Floor, cervical 9 of the same Rapetosaurus individual, from Curry Rogers (2009:fig. 9). The neural spine is a completely different shape from that of C11, but that is presumably mostly due to damage. One of the interesting things here is the apparent lack of pneumatic foramina in the centrum. They’re there somewhere: Curry Rogers (2009:1054) writes “In cervical vertebrae 9, 11, and 12, the centrum bears an elongate shallow pneumatic fossa with two anterior pneumatic foramina surrounded by sharp, lip-like boundaries.” But they are hard to make out! 

CurryRogers2009-rapetosaurus-fig9-C9

The meta-oddity here is that the cervicals of the four titanosaur genera pictures here are all so different from each other. What does this mean?

Probably only that Titanosauria is a huge, disparate, long-lived clade that encompasses far more morphological variation than (say) Diplodocidae. It’s a truism that we don’t, even now, really have a handle on titanosaur phylogeny — every new study that comes out seems to recover a dramatically different topology — so our perception of the clade is really as a big undifferentiated blob. In contrast, the division of Diplodocoidea into Rebbachisaurids, Dicraeosaurids and Diplodocids (plus some odds and ends) is nicely established and easy to think about.

So. Lots of work to be done on titanosaurs.

References

Mounted Alamosaurus in Dallas 1

Next week I’m going to visit the Perot Museum of Nature and Science in Dallas, Texas, to see their big Alamosaurus (these photos were kindly provided by Ron Tykoski of the Perot Museum, with permission to post). See that sweet string of cervical vertebrae in front of the mounted skeleton? A photo of those same vertebrae when they were still in the ground was featured in the post “How big was Alamosaurus?” three and a half years ago. Happily now they are out of the ground, prepped, and on display, and Tony Fiorillo and Ron Tykoski are working on getting them and some other new Alamosaurus material described.

Mounted Alamosaurus in Dallas 2

Here’s another view of that mount. You may be wondering, first, how legit is it, and second, how big is it? Happily, I have answers for you. In email messages with permission to cite, Ron Tykoski wrote,

The Alamosaurus skeletal mount by RCI  in the photos is based upon scaling the Smithsonian and UT Austin material to match the size of our cervicals here in Dallas.  There were enough overlapping parts between the pieces at the three institutions to get the proportions pretty nicely supported.

I ran across your SV-POW thread on ‘How big was Alamosaurus?’ back when you first posted it in ‘09.  You ought to be pleased to know that you came remarkably close to the eventual size of the skeleton we wound up with.  The full skeleton RCI generated (again, based off scaling to the Dallas verts) is 84ft long, about 16ft at the shoulder (I dropped a tape measure from the 1st dorsal neural spine to the floor during skeleton construction and got 480cm-490cm), and a neck + head of about 25ft.  The overall length and neck length were provided by RCI after fabrication and assembly.   That shoulder height is a bit suspect though based on the positioning of the pectoral girdle in the mount, relative to the ribcage and vert column.   I think the head currently is posed about 25ft or so off the floor, but I can’t verify that (I didn’t get into the scissor-lift to check that at the time).  This skeleton actually played a role in determining the size of the hall in which it is installed.  We decided early in the planning phase for the building that this skeleton would be the centerpiece for the hall.  As a result, the ceilings for this floor had to be made extra-high, and the mid-room support pillars designed out to accommodate the skeleton and still clear all the HVAC, sprinkler heads, and other necessities.

That’s all pretty fantastic–both that we have enough of Alamosaurus to do a pretty rigorous full skeletal mount, and that the beast was legitimately pretty darned big. Ron goes on:

One correction to the story on SV-POW, the Dallas cervical series consists of only 9 verts, not 10.  There may have been frags or something that made folks think there was a 10th at the anterior end of the series when first found, but I’ve never seen evidence of it in our collection.  This may be supported by the fact that the verts were given letter designations in the field (that we still use), and are identified as verts B through J, from anterior to posterior.

I later learned from Tony Fiorillo that the vertebrae were labelled B through J in the field in case anything anterior to B turned up, but nothing did, so the ‘A’ placeholder went unused. That reminds me of the search in the mid-1800s for the hypothetical planet Vulcan (not the one you’re thinking of) between Mercury and the Sun, which I bring up for no reasons other than that hypothetical planets are cool, and if you’re exploring, it’s worth keeping an open mind about what might yet turn up.

There’s more to say about the size of Alamosaurus–we haven’t even covered the big material described by Fowler and Sullivan (2011) yet–but I’m not going to say a whole lot right now, since I’m going to see the Big Bend material in Dallas in just a few days. Watch this space.

Reference

Fowler, D.W. and Sullivan, R.M. 2011. The first giant titanosaurian sauropod from the Upper Cretaceous of North America. Acta Palaeontologica Polonica 56 (4): 685–690.

Trust me, you want to click for the full effect.

Trust me, you want to click for the full effect.

This post is just an excuse for me to show off Brian Engh’s entry for the All Yesterdays contest (book here, contest–now closed–here). The title is a reference to this post, by virtue of which I fancy myself at least a spear-carrier in what I will grandly refer to as the All Yesterdays Movement.

Oddly enough, I don’t have a ton to say about this; I think Brian has already explained the thinking behind the piece sufficiently on his own blog. In the brave new world of integumentarily enhanced ornithodirans, these diamantinasaurs are certainly interesting but not particularly outlandish (Brian’s already done outlandish). And it’s pretty darned hard to argue that sauropods never went into caves, although I can’t off the top of my head think of any previous spelunking sauropods (I’m not counting Baylene in Disney’s Dinosaur; feel free to refresh my memory of others in the comments). The glowworms are not proven, but biogeographically and stratigraphically plausible, which is probably as good as we’re going to get given the fossilization potential of bioluminescence.

I’m much more excited about this as a piece of art. I got to see a lot of the in-progress sketches and they were wonderful, with some very tight, detailed pencil-work. The danger in investing that kind of effort is that then you’re tempted to show it off, and if I had any worry about the finished piece, it was that it would be over-lit to show off all the details. But it isn’t. I can tell you from seeing the pencil sketches that the detail went all the way down, but Brian was brave enough to let some of that go, especially on the animals’ legs, to get the lighting effect right. My favorite touches are the reflections in the water, and the fallen pillar in the foreground–toppled by a previous visitor, perhaps–with new mineral deposits already forming on it.

All in all, it takes me back to the best paleoart from my childhood, which made me think, “Wow, these were not monsters or aliens, they were real animals, as real, and as mundane in their own worlds, as deer and coyotes and jackrabbits.” * **

And that’s pretty cool. What do you think?

———-

* Okay, maybe not  in those exact words. I am translating a feeling I had when I was nine through 28 years of subsequent experience and vocabulary expansion.

** My major discovery in the last two decades is that deer and coyotes and jackrabbits are just as exotic as dinosaurs, if only you learn to really see them. And before Mike jumps me for saying that, I said ‘just as exotic’, not ‘just as awesome‘.

UPDATE the next day

If you thought the glowworms were unrealistic–and at least one commenter did–check these out (borrowed from here, pointed out by Brian):

NZ121877D6

NZ121864D6

 

That’s game, set, and match on the glowworm issue.

I was cruising the monographs the other night, looking for new ideas, when the humerus of Opisthocoelicaudia stopped me dead in my tracks. I think you’ll agree it is an arresting sight:

Opisthocoelicaudia right humerus in lateral, anterior, medial, and posterior views, from Borsuk-Bialynicka (1977: figure 7)

Opisthocoelicaudia right humerus in medial, anterior, lateral, and posterior views, from Borsuk-Bialynicka (1977: figure 7)

I’d seen it before, but somehow I had never grokked its grotesque fatness. I mean, damn, Opisthocoelicaudia, you really let yourself go. Especially compared to the slenderness and grace of this juvenile Alamosaurus humerus:

Alamosaurus left humerus in anterior and posterior views, from Lehman and Coulson (2002: figure 7).

Alamosaurus left humerus in anterior and posterior views, from Lehman and Coulson (2002: figure 7).

Now, I realize that part of the slenderness of this Alamosaurus humerus might be because it’s a juvenile–other alamosaur humeri are a bit more robust–but it’s still a striking contrast. I couldn’t help but superimpose them, scaled to the same midshaft width:

Alamosaurus and Opisthocoelicaudia humeri superimposed

I flipped the Alamosaurus humerus left-to-right to match that astonishing lump of Opisthocoelicaudia. The result reminds me of one of Abrell and Thompson’s Actual Facts:

If you put Woodrow Wilson inside William Howard Taft, he would have stuck out by a good 18 inches.

None of that probably signifies anything more than that I am easily amused. And also,  Opisthocoelicaudia is Just Plain Wrong. You hear me, Opisthocoelicaudia? Don’t make me make you cry mayonnaise!

References

You may remember this:

Rapetosaurus mount at Field Museum

…which I used to make this:

Rapetosaurus skeleton silhouette

…and then this:

Rapetosaurus skeleton silhouette - high neck

The middle image is just the skeleton from the top photo cut out from the background and dropped to black using ‘Levels’ in GIMP, with the chevrons scooted up to close the gap imposed by the mounting bar.

The bottom image is the same thing tweaked a bit to repose the skeleton and get rid of some perspective distortion on the limbs. The limb posture is an attempt to reproduce an elephant step cycle from Muybridge.

That neck is wacky. Maybe not as wrong as Omeisaurus, but pretty darned wrong. As I mentioned in the previous Rapetosaurus skeleton post, the cervicals are taller than the dorsals, which is opposite the condition in every other sauropod I’ve seen. All in all, I find the reposed Rapetosaurus disturbingly horse-like. And oddly slender through the torso, dorsoventrally at least. The dorsal ribs look short in these lateral views because they’re mounted at a very odd, laterally-projecting angle that I think is probably not correct. But the ventral body profile still had to meet the distal ends of the pubes and ischia, which really can’t go anywhere without disarticulating the ilia from the sacrum (and cranking the pubes down would only force the distal ends of the ilia up, even closer to the tail–the animal still had to run its digestive and urogenital pipes through there!). So the torso was deeper than these ribs suggest, but it was still not super-deep. Contrast this with Opisthocoelicaudia, where the pubes stick down past the knees–now that was a tubby sauropod. Then again, Alamosaurus has been reconstructed with a similarly compact torso compared to its limbs–see the sketched-in ventral body profile in the skeletal recon from Lehman and Coulson (2002: figure 11).

I intend to post more photos of the mount, including some close-ups and some from different angles, and talk more about how the animal was shaped in life. And hopefully soon, because history has shown that if I don’t strike while the iron is hot, it might be a while before I get back to it. For example, I originally intended this post to follow the last Rapetosaurus skeleton post by  about a week. So much for that!

Like everything else we post, these images are CC BY, so feel free to take them and use them. If you use them for the basis of anything cool, like a muscle reconstruction or life restoration, let us know and we’ll probably blog it.

Caudal pneumaticity in saltasaurines. Cerda et al. (2012: fig. 1).

Earlier this month I was amazed to see the new paper by Cerda et al. (2012), “Extreme postcranial pneumaticity in sauropod dinosaurs from South America.” The title is dramatic, but the paper delivers the promised extremeness in spades. Almost every figure in the paper is a gobsmacker, starting with Figure 1, which shows pneumatic foramina and cavities in the middle and even distal caudals of Rocasaurus, Neuquensaurus, and Saltasaurus. This is most welcome. Since the 1990s there have been reports of saltasaurs with “spongy bone” in their tail vertebrae, but it hasn’t been clear until now whether that “spongy bone” meant pneumatic air cells or just normal marrow-filled trabecular bone. The answer is air cells, loads of ‘em, way farther down the tail than I expected.

Caudal pneumaticity in diplodocines. Top, transverse cross-section through an anterior caudal of Tornieria, from Janensch (1947: fig. 9). Bottom, caudals of Diplodocus, from Osborn (1899: fig. 13).

Here’s why this is awesome. Lateral fossae occur in the proximal caudals of lots of neosauropods, maybe most, but only a few taxa go in for really invasive caudal pneumaticity with big internal chambers. In fact, the only other sauropod clade with such extensive pneumaticity so far down the tail are the diplodocines, including Diplodocus, Barosaurus, and Tornieria. But they do things differently, with BIG, “pleurocoel”-type foramina on the lateral surfaces of the centra, leading to BIG–but simple–camerae inside, and vertebral cross-sections that look like I-beams. In contrast, the saltasaurines have numerous small foramina on the centrum and neural arch that lead to complexes of small pneumatic camellae, giving their vertebrae honeycomb cross-sections. So caudal pneumaticity in diplodocines and saltsaurines is convergent in its presence and extent but clade-specific in its development. Pneumaticity doesn’t get much cooler than that.

Pneumatic ilia in saltasaurines. Cerda et al. (2012: fig. 3).

But it does get a little cooler. Because the stuff in the rest of the paper is even more mind-blowing. Cerda et al. (2012) go on to describe and illustrate–compellingly, with photos–pneumatic cavities in the ilia, scapulae, and coracoids of saltasaurines. And, crucially, these cavities are connected to the outside by pneumatic foramina. This is important. Chambers have been reported in the ilia of several sauropods, mostly somphospondyls but also in the diplodocoid Amazonsaurus. But it hasn’t been clear until now whether those chambers connected to the outside. No patent foramen, no pneumaticity. It seemed unlikely that these sauropods had big marrow-filled vacuities in their ilia–as far as I know, all of the non-pneumatic ilia out there in Tetrapoda are filled with trabecular bone, and big open marrow spaces only occur in the long bones of the limbs. And, as I noted in my 2009 paper, the phylogenetic distribution of iliac chambers is consistent with pneumaticity, in that the chambers are only found in those sauropods that already have sacral pneumaticity (showing that pneumatic diverticula were already loose in their rear ends). But it’s nice to have confirmation.

So, the pneumatic ilia in Rocasaurus, Neuquensaurus, and Saltasaurus are cool because they suggest that all the other big chambers in sauropod ilia were pneumatic as well. And for those of you keeping score at home, that’s another parallel acquisition in Diplodocoidea and Somphospondyli (given the apparent absence of iliac chambers in Camarasaurus and the brachiosaurids, although maybe we should bust open a few brachiosaur ilia just to be sure*).

* I kid, I kid.**

** Seriously, though, if you “drop” one and find some chambers, call me!

Pectoral pneumaticity in saltasaurines. Cerda et al. (2012: fig. 2).

But that’s not all. The possibility of pneumatic ilia has been floating around for a while now, and most of us who were aware of the iliac chambers in sauropods probably assumed that eventually someone would find the specimens that would show that they were pneumatic. At least, that was my assumption, and as far as I know no-one ever floated an alternative hypothesis to explain the chambers. But I certainly did not expect pneumaticity in the shoulder girdle. And yet there they are: chambers with associated foramina in the scap and coracoid of Saltasaurus and in the coracoid of Neuquensaurus. Wacky. And extremely important, because this is the first evidence that sauropods had clavicular air sacs like those of theropods and pterosaurs. So either all three clades evolved a shedload of air sacs independently, or the basic layout of the avian respiratory system was already present in the ancestral ornithodiran. I know where I’d put my money.

There’s loads more interesting stuff to talk about, like the fact that the ultra-pneumatic saltasaurines are among the smallest sauropods, or the way that fossae and camerae are evolutionary antecedent to camellae in the vertebrae of sauropods, so maybe we should start looking for fossae and camerae in the girdle bones of other sauropods, or further macroevolutionary parallels in the evolution of pneumaticity in pterosaurs, sauropods, and theropods. Each one of those things could be a blog post or maybe a whole dissertation. But my mind is already thoroughly blown. I’m going to go lie down for a while. Congratulations to Cerda et al. on what is probably the most important paper ever written on sauropod pneumaticity.

References

  • Cerda, I.A., Salgado, L., and Powell, J.E. 2012. Extreme postcranial pneumaticity in sauropod dinosaurs from South America. Palaeontologische Zeitschrift. DOI 10.1007/s12542-012-0140-6
  • Janensch, W. 1947. Pneumatizitat bei Wirbeln von Sauropoden und anderen Saurischien. Palaeontographica, Supplement 7, 3:1–25.
  • Osborn, H. F. 1899. A skeleton of Diplodocus. Memoirs of the American Museum of Natural History 1:191–214.

Thanks to the kind offices of the folks at the Field Museum, especially Fossil Vertebrates collection manager Bill Simpson, on Wednesday I got to hop the fence and spend some quality time with FMNH PR 2209, the mounted holotype specimen of Rapetosaurus krausei. I took a tape measure with me, to get some dimensions from the mounted skeleton. Of course I have the detailed descriptive paper (Curry-Rogers 2009), but mounted skeletons are three-dimensional objects and it is often surprisingly difficult to get a sense of a how a skeleton goes together in three dimensions from pictures and measurements of the individual elements. And if these dimensions are not precisely those of the animal in life, because of assumptions made during mounting–concerning, say, cartilage thickness between bones, or the angles of the ribs–at least they’re a starting point for understanding the whole-body proportions of Rapetosaurus.

This is valuable because AFAIK this specimen is the only mounted titanosaur in North America, and maybe the only one outside of South America and China. [UPDATE: Alert commenters pointed out that I forgot about the Opisthocoelicaudia in Warsaw, which is almost entirely real, and the Argentinosaurus in Georgia, which is almost entirely fake.] And because Rapetosaurus is far out, man. ALL of the neural arches are unfused, even in the distal caudals–even the Arundel Astrodon (formerly Pleurocoelus) material has fused arches in the distal caudals (Wedel et al. 2000: fig. 15). So it’s a very young juvenile, but the neck is already more than twice the length of the body. I say ‘already’ because there is pretty good evidence that the cervical vertebrae grew proportionally longer over the course of ontogeny in at least some sauropods (Wedel et al. 2000:368-369). The neck is 336 cm long, and the femora are 69 cm long. If we isometrically scaled this animal up to have a 2-meter femur, the neck would be 10 meters long, without any such ontogenetic telescoping of the vertebrae. The implications of this for possible neck lengths in the supergiant titanosaurs are pretty darned interesting. The vertebrae of Rapetosaurus don’t really look anything like those of Argentinosaurus. Nevertheless, a sauropod with an Argentinosaurus-sized femur (2.5 meters for the largest known) and Rapetosaurus proportions would have a 12-meter neck–again, that’s assuming this very young Rapetosaurus already has adult proportions, when in fact it may be ontogenetically short-necked (now there’s a thought). In Apatosaurus and Camarasaurus, the cervicals grew in proportional length (i.e., relative to diameter) by 30-50% over ontogeny, but that’s starting from tiny baby vertebrae. The Rapetosaurus vertebrae are already very long, proportionally, but it is interesting to consider the possibilities that they might have been even longer in adults, and that that scaling might have been shared with other titanosaurs.

The tail in this mount is oddly short. Only about every third vertebra is real, with the rest sculpted, so the tail length inevitably depends on how many intermediary vertebrae were added. But unless there are a LOT of missing vertebrae, it’s probably not far off. I can tell you that when I first saw the mount I looked at the tail and said, “No way”. But up close, seeing the real vertebrae and the interspersed intermediates, it looked pretty reasonable, in part because the individual caudal vertebrae are proportionally short. This is one of those things where we may just have to wait for more and better material–although that might be a long wait, because this skeleton is already freakin’ gorgeous. For someone who is used to dealing with hideously incomplete and groadily distorted fossils, this Rapetosaurus material is just mouth-wateringly beautiful.

There’s loads more weird stuff to talk about, like how the cervical vertebrae are taller than the dorsals, which is opposite the condition in every other sauropod I’ve gotten to look at, and the shape of the ilium, and the conformation of the rib cage, but those will all have to wait for future posts. This one is already much longer than I intended it to be (standard).

For the curious, here are all of my measurements. Neck length, dorsal length, etc. are lengths of those sections of the column as mounted–that is, including both the vertebrae and the spaces between them. I haven’t compared any of these to the published measurements, these are straight from the tape measure to my notebook to you. I’m giving them in mm, because that’s what I naturally think in, but they’re all rounded to the nearest cm because given my methods–hand-holding a physical tape measure up next to a bone while I crouch contorted under a fragile mounted skeleton–giving measurements to the nearest mm would be illusory precision.

  • Skull length: 290
  • Neck length: 3360
  • Dorsal length: 1210
  • Sacrum length: 480
  • Tail length: 1720
  • Total length of skeleton, snout to tip of tail (sum of above): 7060
  • Glenoid height (ground to top of socket): L – 1110 (forefoot off floor by a few cm), R – 1080
  • Acetabular height (ground to top of socket): 1320 on both sides
  • Max height of body (ground to top of 5th sacral spine): 1630
  • Gleno-acetabular distance: L – 1500, R – 1440
  • Width across acetabula: 440 between weight-bearing centers, 470 to outer margins of ilia
  • With across glenoids (at bottom of scap-coracoid joints): 710
  • Femur length: 690 on both sides
  • Tib/fib length: 470 on both sides
  • Vertical height of foot: L – 90, R – 120 (different poses)
  • Humerus length: L – 530, R – 500
  • Radius/ulna length (between articular surfaces, not including olecranons): L – 370, R – 360
  • Metacarpus length (MT3): 190 on both sides

References

[This is a guest post by frequent commenter Heinrich Mallison.  Heinrich is maybe best known to SV-POW! readers for his work on digital modelling of sauropodomorphs, though that may change now that his paper on sauropod rearing mechanics is out.  Read on ...]

Maybe this post should have been titled “How sauropods breathed, ate, and farted”. Or maybe not. But breathing, eating and fermenting the food will play an important role.

Last week held a special pleasure for me. I spent it in New York, digitizing sauropods bones in the American Museum of Natural History’s Big Bone Room. Treasure trove that this room is, the museum still held something even better: the opening of a new special exhibit titled The World’s Largest Dinosaurs. While all such exhibits are of general interest to me, this one is special. Mark Norell, famous palaeontologist and curator at the AMNH, had a co-curator for this exhibit, Martin Sander of Bonn University, who is the head and speaker of the German Research Foundation Research Unit FOR 533 “Sauropod Biology”. As a member of FOR 533, and having received funding for both my PhD work and my first post-doc project, I am obviously somewhat biased, so please take this into account when you read this report.

The exhibition does not show a large amount of sauropods material. Not that it wouldn’t make for a nice exhibit, as the AMNH’s Hall of Saurischian Dinosaurs doesn’t really have that many sauropods (one Apatosaurus mount, to be exact, with a mashed up Barosaurus vertebral column half-hidden away and a wonderful but obviously depressed “prosauropod”, my old friend Plateosaurus, thrown in to make up a bit for the many, many stinkin’ theropod specimens). But instead of showcasing some of the usually hidden-away bones of the AMNH collection (and believe me, there is some wonderful stuff there), it rather focuses on those parts of the animal that are usually missing: the soft tissues. “How did sauropods get so big?”, or, reversing the question: “Why did and does no other group of terrestrial vertebrates reach such gigantic body sizes?” These were the questions our research group has been busily investigating for the last six years, and the answers to these question are what the exhibit now tries to communicate to the public. And it does so quite successfully!

The centerpiece of the AMNH exhibit: the belly of Mama Mamenchisaurus.

The centrepiece is a full-sized, fleshed out model of a sauropod (Mamenchisaurus hochuanensis), but on one side the skin and superficial musculature has been cut away. The visitor can see the neck vertebrae, the trachea, the carotid artery, and the ribcage. And the ribcage is also a projection area, on which a video is played that shows the internal organs and how they work.

With a voice-over that explains the actions in simple terms, the principle of the avian-style unidirectional lung and the air sacs is explained (albeit with a small error, as lung physiologist and FOR 533 member Steve Perry was quick to point out – the AMNH has promised to fix things), as well as the basic principles of sauropod reproduction (high number of offspring). Many things are not said or shown here, which is a good thing as it allows for the normal short attention span of the average museum visitor for one piece of exhibit. Instead, interesting stuff like how much fodder a sauropod needed per day (or even per hour), a comparison of a sauropod’s and an elephant’s heart, and of a giraffe’s and a sauropod’s neck vertebra (wow, how light the sauropod one is!) are explored at small science stations spread around the room. I won’t go into a detailed description here, you can find that elsewhere on the web. The AMNH did a blogger’s preview a while ago, and invited the press for a press conference and walk-through of the exhibit with the chance to interview the scientists present on Wednesday, so much info has already been plastered all over the web. Instead, I’ll just show you some pics and talk a bit about the concept of the exhibition, and how various issues were handled that can make or break a show.

One thing is how to catch the attention of visitors and direct it to the content of the exhibit. You don’t want people just going “aw, sh*t! That is one HUGE bone/animal!” and wandering off into the next room. If you want to educate them (and that, may I remind you, is the central purpose of a museum exhibit), you need to get them interested in stuff. Get them to read texts, look at stuff (not just let their eyes wander across it for a few seconds), try to get their brains going. The sauropod exhibit manages this by, first of all, being behind a closed door you can’t see through. Usually, the AMNH halls are accessible either through an open doorway, or in a few cases through glass doors. Secondly, the exhibit, especially the rather confined area you enter first, is dark. Very dark. Again a marked contrast to the AMNH’s usually well-lit halls. Just a few plants greet the visitor, and it takes a second to adjust to the dark – enough time to look around a bit and notice the neck and head of Argentinosaurus (fleshed out model) above.

My esteemed colleague Vivian Allen from Royal Veterinary College London going "Aw, sh*t! That is one HUGE sauropod!"

Next, the visitor is channeled along, with only a very few specimens to catch his attention. Well done, because these few pieces (sauropod leg, Komodo dragon skeleton, human skeleton, etc.) focus on getting the main message across (sauropods = way larger than everything else), aided by the largest animals (or their silhouettes) or various groups painted on the wall. Only once the message has been driven home, as I could detect from the comments I overheard, are the visitors released into the main area that contains the sauropod model and the various detail exhibits around it.

The next thing is giving people time to check things out. If you herd them too much, they will get driven along by the masses. That’s why the larger, opener area around the sauropod model and the smaller bits around it works so well: people can sit down to see the projected videos on the sauropod belly, or they can drift around from one specimen or science station to the next.

The stations are not just glass cabinets with some bones in them. Instead, at many of them you can DO things. One allows you to measure either an adult or baby sauropod femur or your own, and then calculate how heavy a sauropod of that size was. At another you can pump a sauropod’s and an elephant’s lung. One I liked very much simply had an unpainted sauropod model, and two sets each (adult and children height) of oculars. One showed a colorful “show-off” version, the other a “camouflage” one. “Which one is true? We don’t know!” is how I’d paraphrase the text that goes with it. One that innocently hides in the corner is among the most impressive: a 5 ½ ft cube (1.7 m, for the civilized) made from Plexiglas filled with sauropod food. A serving sufficient for one day! On it, also, the various plant groups available in the Mesozoic were rated for various factors, getting an easily understood rating in stars. That’s another big thing: make things easily understandable, visualize them!

Yummy! 100% Recommended Daily Value for your average sauropod.

With all these things well done, there remains only one more thing: make things fun for kids! And the AMNH did just that by adding a kids’ dinosaur dig. OK, it is one of those cheesy things where you use brushes and stuff to brush sand off fossils (cast), but it was done well enough that kids lined up like there was no tomorrow.

Overall, the exhibit gets two big thumbs up from me. If you make it to NY while it is on, or to any of its future stations, go see it! However, as FOR 533 member Steve Perry was quick to point out: if you’re in it only for the size, you’ll be disappointed! Aside from a few isolated bones, not much of the largest dinosaurs (Argentinosaurus and Amphicoelias) is to be seen in bone. It is the biological details that matter!  But don’t get me started about the tail musculature, especially the caudofemoralis, of the big model.

And then, there is the other thing about it that is closely tied to shameless self-promotion: the AMNH did not produce a catalogue or anything similar. Instead, the latest book from the “Life of the Past” series (Editor: James Farlow) of Indiana University Press was presented at the press conference. The lucky reporters all even got a free copy! The title is Biology of the Sauropod Dinosaurs: Understanding the Life of Giants, edited by N. Klein, K. Remes, C. T. Gee and P. M. Sander. And by now, I am sure, you have figured out who the authors are … It is intended to be a summary of the research findings of the first (and part of the second) funding period of FOR 533, and yours truly has two chapters in it. The first doesn’t really give much new information; most is already contained in my two papers here and here. The second, however, presents novel research that didn’t make it into the AMNH exhibit. But hey, why spoil the surprise – go and buy our book!) Overall, it is quite a technical book, so laypeople beware, but we did try to make the research as accessible as possible while retaining a high standard. For the even more technically minded there is the summary of our research group’s work (which cost the DFG ~€6.000.000) to be found in Sander et al. 2010. However, reading that paper is not half as much fun as the book, or the exhibit.

References

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