AMNH 460 left anterolateral view

Apatosaurines on the brain right now.

I’ve been thinking about the question raised by Jerry Alpern, a volunteer tour guide at the AMNH, regarding the recent Tschopp et al. (2015) diplodocid phylogeny. Namely, if AMNH 460 is now an indeterminate apatosaurine, pending further study, what should the museum and its docents tell the public about it?

Geez, Apatosaurus, it’s not like we’re married!

I think it’s a genuinely hard problem because scientific and lay perspectives on facts and hypotheses often differ. If I say, “This animal is Apatosaurus“, that’s a fact if I’m talking about YPM 1860, the genoholotype of Apatosaurus ajax; it would continue to be a fact even if Apatosaurus was sunk into another genus (as Brontosaurus was for so long). We might call that specimen something else, but there would always be a footnote pointing out that it was still the holotype of A. ajax, even if the A. part was at least temporarily defunct – the scientific equivalent of a maiden name.* For every other specimen in the world, the statement, “This animal is Apatosaurus” is a hypothesis about relatedness, subject to further revision.

* This is going to sound kinda horrible, but when one partner in a marriage takes the other’s surname, that’s a nomenclatural hypothesis about the future of the relationship.

Apatosaurine cervicals are the best cervicals.

Apatosaurine cervicals are the best cervicals.

Fuzzy science

Things that look fairly solid and unchanging from a distance – specifically, from the perspective of the public – often (always?) turn out to be fairly fuzzy or even arbitrary upon closer inspection. Like what is Apatosaurus (beyond the holotype, I mean) – or indeed, what is a planet.** There is no absolute truth to quest for here, only categories and hypotheses that scientists have made up so that we can have constructive conversations about the crazy spectrum of possibilities that nature presents us. We try to ground those categories and hypotheses in evidence, but there will always be edge cases, and words will always break down if you push them too hard. Those of us who work on the ragged frontier of science tend to be fairly comfortable with these inescapable uncertainties, but I can understand why people might get frustrated when they just want to know what the damned dinosaur is called.

** Triton, the largest body orbiting Neptune, is almost certainly a captured Kuiper Belt object, and it’s bigger than Pluto. Moon or planet? Probably best to say a former dwarf planet currently operating as a satellite of Neptune – but that’s a mouthful (and a mindful, if you stop to think about it), not a short, convenient, easily-digestible label. Any short label is going to omit important information. This is related to the problem of paper title length – below some threshold, making something shorter means making it incomplete.

What I would say

I suppose the short version that is most faithful to the Tschopp et al. results is:

This skeleton (AMNH 460) might be Apatosaurus or Brontosaurus or a third, new thing – scientists aren’t sure yet.

A reasonable follow-up sentence – and an answer to the inevitable “Why not?” – would be:

They have to look at 477 anatomical details for lots of skeletons and weigh all the evidence, and that takes time.

Personally, if I was talking to museum visitors I would lean in conspiratorially and say:

If you want to call it Apatosaurus or Brontosaurus, go ahead – those are both ‘live’ hypotheses, and even the world’s experts on this problem can’t tell you that you’re guessing the wrong way – at least not yet.

And if there was a kid in the group, I’d add:

Maybe you’ll be the one to figure it out!

What would you say?

My neck is fat.

My neck is fat.

P.S. I wouldn’t change the signage. It could still turn out to be Apatosaurus, and the Tschopp et al. results do not lend themselves to easy label-ification.

P.P.S. With some modification for taxonomy, all of this applies to the Field Museum diplodocid FMNH P25112 as well.

Reference

Tschopp E, Mateus O, Benson RBJ. (2015) A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda) PeerJ3:e857 https://dx.doi.org/10.7717/peerj.857

Look on my works, ye mighty, and despair!

DSCN0476

[Giraffatitan brancai paralectotype MB.R.2181 (formerly HMN S II), mounted skeleton in left anteroventrolateral view. Presacral vertebrae sculpted, skull scaled and 3d-printed from specimen T1. Round the decay of that colossal wreck, boundless and bare, the lone and level sands stretch far away.]

Brontosaurus, the animal formerly known as Apatosaurus, the animal formerly known as Brontosaurus.

YPM 1980: Brontosaurus excelsus, the animal formerly known as Apatosaurus excelsus, the animal formerly known as Brontosaurus excelsus.

Today is a good day for sauropod science. Since we’re not getting this up until the afternoon, you’ve probably already seen that Emanuel Tschopp and colleagues have published a monstrous specimen-level phylogenetic analysis of Diplodocidae and, among other things, resurrected Brontosaurus as a valid genus. The paper is in PeerJ so you can read it for free (here).

I’ve already been pinged by lots of folks asking for my thoughts on this. I know that the return of Brontosaurus is what’s going to catapult this paper into the spotlight, but I hope what everyone takes away from it is just what a thorough piece of work it is. I’ve never seen so many phylogenetic characters illustrated so well. It sets a new standard, and anyone who wants to overturn this had better roll up their sleeves and bring a boatload of data. I’m also very, very happy that it’s open-access so everyone in the world can see it, use it, question it, tear it apart or build on it. Getting Brontosaurus back is just gravy. Although, being pro-brontosaur enough to have named a dinosaur in honor of Brontosaurus, I’m also pretty happy about that. If you need a quick guide to who’s who now, A. ajax and A. louisae are still Apatosaurus, and B. excelsus, B. yahnahpin (formerly Eobrontosaurus), and B. parvus (originally Elosaurus) are all Brontosaurus. For more details, go read the paper.

A louisae from Wikipedia - full

Apatosaurus lousiae CM 3018: still Apatosaurus. Photo from Wikipedia.

My personal feelings aside, a lot of people are asking how solid is this generic re-separation. I haven’t read the entire paper yet – it’s 299 pages long, for crying out loud – but the separation of Brontosaurus and Apatosaurus seems solid enough. Tschopp et al. didn’t do it lightly, they justify their decision in detail. I don’t hold with the idea that just because two taxa are sisters, means that they cannot be separated generically. As usual in phylogenetic taxonomy, it comes down to what we decide as a community constitutes “diagnosably distinct”. Tschopp et al. have actually put some thought into what that might mean here, and whether you agree with them or not, they’ve at least made all of their evidence and reasoning explicit. That’s both an opportunity and a challenge for critics: an opportunity to pin down exactly where and why you may disagree, and a challenge to do exactly that. You can’t just sit back and say, “I think the analysis is flawed” or “I wouldn’t have coded that character that way” (well, you can, but if that’s all you say, no-one is obliged to take that kind of lazy, drive-by criticism seriously). There are 477 characters here, most of them illustrated, for 81 OTUs, and a lot of post-hoc discussion of the results. So whether you agree with the authors or not, in whole or in part, both fans and critics should dig in and build on this work. Is it the last word on diplodocid taxonomy? Of course not. But it does move the field forward significantly, and the Tschopp et al. should be applauded for that.

There’s a lot more in there than just bringing back Brontosaurus. “Diplodocus” hayi is elevated to its own genus, Galeamopus. Neither of those things are super surprising. There have been rumors since the 90s at least that Brontosaurus might be coming back, and everyone has known for a while that D. hayi was a bit wonky. I was also not surprised to see Australodocus returned to Diplodocidae – when I saw the type material in 2011, it looked diplodocid to me (based on some characters I’ll have to unpack in some other post). More surprising to me are the sinking of Dinheirosaurus into Supersaurus, the finding that Tornieria is not particularly close to Diplodocus, and the uncertain positions of AMNH 460, the American Museum mount, which is an indeterminate apatosaurine pending further study, of FMNH 25112, the Field Museum “Apatosaurus”, which might not even be an apatosaurine at all(!). In several cases, Tschopp et al. come right out and say that X is going to need further study, so if you want to work on sauropods and you’re stuck for project ideas, go see what needs doing.

AMNH mounted Apatosaurus with Taylor for scale

AMNH 460: we don’t know who this is anymore.

As I was scanning the paper again while composing the last paragraph, I almost fell down the rabbit hole. So much interesting stuff in this paper. Even if all you care about is morphology, the hundred or so figures illustrating the phylogenetic characters ought to keep you happy for a very long time. I look forward to reading through the vertebral characters in detail and seeing what I’ve been missing all these years.

I’m contractually obliged to point out that the authors chose to publish the complete peer-review history of the paper, so you can see what the editor (Andy Farke) and reviewers had to say. As always, I think this transparency (and credit for the reviewers) is great for science, and I can’t wait until it’s the norm at more journals.

FMNH 25112 formerly Apatosaurus

FMNH 25112: what even IS that thing?

In addition to the paper, there’s also an interview with lead author Emanuel Tschopp on the PeerJ blog, and a nice shout-out for SV-POW!

Parting shot: why did Tschopp et al. get different results than anyone had previously? Because they used more specimens and more taxa – more data full stop. That’s also why their paper warrants serious consideration. It’s serious work. Let’s go stand on their shoulders.

Reference

Tschopp E, Mateus O, Benson RBJ. (2015) A specimen-level phylogenetic analysis and taxonomic revision of Diplodocidae (Dinosauria, Sauropoda) PeerJ 3:e857 https://dx.doi.org/10.7717/peerj.857

In 2012, Matt and I spent a week in New York, mostly working at the AMNH on Apatosaurusminimus and a few other specimens that caught our eye. But we were able to spend a day at the Yale Peabody Museum up in New Haven, Connecticut, to check out the caudal pneumaticity in the mounted Apatosaurus (= “Brontosaurus“) excelsus, YPM 1980, and the bizarrely broad cervicals of the Barosaurus lentus holotype YPM 429.

While we there, it would have been churlish not to pay some attention to the glorious and justly famous Age of Reptiles mural, painted by Rudolph F. Zallinger from 1944-1947.

So here it is, with the Brontosaurus neck for scale:

IMG_0501-zallinger-mural

Click through for high resolution (3552 × 2664).

And here is a close-up of the most important, charismatic, part of the mural:

IMG_0500-zallinger-mural

Again, click through for high resolution (3552 × 2664).

That’s your lot for now. We’ve long promised a proper photo post of the Brontosaurus mount itself, and I’ll try to get that done soon. For now, it’s just scenery.

A while back, Ben Miller reminded me that when I posted about the old Yale “Brontosaurus” skull, I promised:

So how did the YPM come to make such a monstrosity? What was it based on? Tune in next time for the surprising details!

I told him at the time that I’d soon get around to writing a post. But before I did, he wrote a post on this himself: Bully for Camarasaurus. And it’s excellent. Go and read it!

I don’t have a lot to add to what Ben has written, except regarding this:

What Marsh had instead [when restoring the skull for his 1891 “Brontosaurus” reconstruction] were a few fragmentary bits of Camarasaurus cranial material, plus a snout and jaw (USNM 5730) now considered to be Brachiosaurus.

Here’s what Marsh came up with:

Marsh1891-plateXVI-Apatosaurus-skull-UNREVERSED

But what of the supposed Brachiosaurus skull that he used as a reference? It was finally described 107 years later by Carpenter and Tidwell (1998), in a paper that helpfully also lays out the history behind it. Here’s how it looks:

CarpenterTidwell1998-fig1

The skull was found by a crew under the supervision of M. P. Felch in the western part of his Quarry 1, Garden Park, Colorado. Felch reported it to O. C. Marsh in a letter of 8 September 1883. It was found by a meter-long cervical vertebra that probably belonged to Brachiosaurus “which was destroyed during attempts to collect it” (McIntosh and Berman 1975:196). [Of course, Felch and Marsh could hardly have been expected to identify this vertebra correctly, as Brachiosaurus would not be discovered and named for another twenty years (Riggs 1903), and the nature of its neck would not become apparent until Janensch (1914) described the related brachiosaurid Giraffatitan (= “Brachiosaurus“) brancai.]

The Felch skull, along with other material from the quarry, was shipped to Marsh at Yale in October of that year, and was initially assigned the specimen number YPM 1986. At that time it was only partially prepared, hence the rather poor resemblance between the restored version above and Marsh’s hypothetical “Brontosaurus” [= Apatosaurus] skull that was based on it.

It’s notable that Holland (1915) was quite certain that this was not a skull of Brontosaurus, and that a Diplodocus-like skull found with the A. louisae holotype belonged to it. It’s worth reading the skull section of his paper to see just how solid his reasoning was. And it’s extraordinary to think that Osborn’s power, all the way over in New York, was so great that he was able to successfully bully Holland, 370 miles away in Pittsburgh, into not putting the evidently correct skull on the Carnegie Museum’s Apatosaurus mount. That mount remained sadly headless until after Holland’s death.

Aaanyway, YPM 1986 was pretty much ignored after Marsh’s abuse of it as a reference for the Brontosaurus reconstruction’s skull. After Marsh’s death in 1899, much of the material collected by Felch was transferred to the Smithsonian (US National Museum of Natural History). The skull was among these specimens, and so was re-catalogued as USNM 5730.

As so often, it was Jack McIntosh who rediscovered this skull and recognised its true affinities. Some time after his tentative identification of the skull as pertaining to Brachiosaurus (presumably on the basis of its resemblance to that of Giraffatitan), Carpenter borrowed the skull, had it more fully prepared, wrote the description, and had a restored model constructed from casts of the preserved elements and models of the missing ones.

Carpenter and Tidwell (1998:fig. 2) also handily showed the restored Felch quarry skull alongside those of other sauropods:

CarpenterTidwell1998-fig2

By re-ordering the top row, we can see what a neat intermediate it is between the skulls of Camarasaurus (left) and Giraffatitan (= “Brachiosaurus” of their usage):

CarpenterTidwell1998-fig2-top-row-reordered

I provisionally accepted USNM 5730 as belonging to Brachiosaurus in my re-evaluation of 2009, and included it in my reconstruction (Taylor 2009:fig. 7):

Taylor (2007: figure 7). Skeletal reconstruction of Brachiosaurus altithorax. White bones represent the elements of the holotype FMNH P 25107. Light grey bones represent material referred to B. altithorax: the Felch Quarry skull USNM 5730, the cervical vertebrae BYU 12866 (C?5) and BYU 12867 (C?10), the "Ultrasauros" scapulocoracoid BYU 9462, the Potter Creek left humerus USNM 21903, left radius and right metacarpal III BYU 4744, and the left metacarpal II OMNH 01138. Dark grey bones modified from Paul's (1988) reconstruction of Giraffatitan brancai. Scale bar equals 2 m.

Taylor (2007: figure 7). Skeletal reconstruction of Brachiosaurus altithorax. White bones represent the elements of the holotype FMNH P 25107. Light grey bones represent material referred to B. altithorax: the Felch Quarry skull USNM 5730, the cervical vertebrae BYU 12866 (C?5) and BYU 12867 (C?10), the “Ultrasauros” scapulocoracoid BYU 9462, the Potter Creek left humerus USNM 21903, left radius and right metacarpal III BYU 4744, and the left metacarpal II OMNH 01138. Dark grey bones modified from Paul’s (1988) reconstruction of Giraffatitan brancai. Scale bar equals 2 m.

But as noted by Carpenter and Tidwell (1998:82), the lack of comparable parts between the Felch skull and the Brachiosaurus holotype (which remains the only definitive Brachiosaurus material) means that the assignment has to remain tentative.

What we really need is a more complete Brachiosaurus specimen: one with both a skull and good postcervical elements that let us refer it definitively to Brachiosaurus altithorax by comparison with the holotype. And heck, while we’re at it, let’s have a specimen with a good neck, too!

The real question remains: how did Marsh, using a brachiosaur skull as his basis, come up with this?

Marsh1891-plateXVI-Apatosaurus-skull-UNREVERSED

 

And stranger still, how someone at the Yale Peabody Museum — we don’t know who — used it, or more likely Marsh’s reconstruction, as a basis for this sculpture:

IMG_0517

 

The Yale mount didn’t go up until 1931 — the last of the Big Four Apatosaurus mounts after the AMNH, Carnegie and Field Museum, which is surprising as it was the first of those specimens to be found. So by the time the skull was sculpted, sauropod skulls were actually reasonably well known. It’s not clear quite how anyone working from a decent reconstruction of, say, a Camarasaurus skull — the one in Osborn and Mook (1921:figure 30), say — could come up with this monster.

The last thing to say is this: it does credit to the YPM that they display this historically important sculpture rather than hiding it away and pretending it never happened. For me, part of the fascination of palaeontology is seeing not just how organisms evolved through prehistory but how ideas evolved through history. It’s great that we can still see important mistakes, alongside their corrections (i.e. the new and lovely skull on the YPM Apatosaurus mount.)

 

References

  • Carpenter, Kenneth, and Virginia Tidwell. 1998. Preliminary description of a Brachiosaurus skull from Felch Quarry 1, Garden Park, Colorado. Modern Geology 23:69-84.
  • Holland, William J. 1915. Heads and tails: a few notes relating to the structure of the sauropod dinosaurs. Annals of the Carnegie Museum 9:273-278.
  • Janensch, Werner. 1914. Ubersicht uber der Wirbeltierfauna der Tendaguru-Schichten nebst einer kurzen Charakterisierung der neu aufgefuhrten Arten von Sauropoden. Archiv fur Biontologie, Berlin III, 1(1):81-110.
  • Marsh, O. C. 1891. Restoration of Triceratops (with plates XV and XVI). American Journal of Science, 3rd series 41(244):339-342.
  • McIntosh, John S., and David, S. Berman. 1975. Description of the palate and lower jaw of the sauropod dinosaur Diplodocus (Reptilia: Saurischia) with remarks on the nature of the skull of Apatosaurus. Journal of Paleontology 49(1):187-199.
  • Osborn, Henry Fairfield, and Charles C. Mook. 1921. Camarasaurus, Amphicoelias and other sauropods of Cope. Memoirs of the American Museum of Natural History, n.s. 3:247-387, and plates LX-LXXXV.
  • Riggs, Elmer S. 1903. Brachiosaurus altithorax, the largest known dinosaur. American Journal of Science 15(4):299-306.
  • Taylor, Michael P. 2009. A re-evaluation of Brachiosaurus altithorax Riggs 1903 (Dinosauria, Sauropoda) and its generic separation from Giraffatitan brancai (Janensch 1914). Journal of Vertebrate Paleontology 29(3):787-806.

 

We all remember Upchurch and Martin’s (2002) description of the Rutland Cetiosaurus, which remains by some distance the best British sauropod specimen in the literature; and the same authors’ (2003) survey of the genus Cetiosaurus. They concluded that nearly all of its many named species are either nomen dubia or misassigned, and that only C. oxoniensis is a valid, diagnosable species.

(Some of) the Cetiosaurus oxoniensis holotype material, on display in the public gallery of the Oxford University Museum of Natural History (OUMNH)

(Some of) the Cetiosaurus oxoniensis holotype material, on display in the public gallery of the Oxford University Museum of Natural History (OUMNH). From left to right: right femur in posterior view, scapula, right humerus in anterior view, tibia and fibula (designations by eyeballing). Above the long bones, some caudal vertebrae.

Accordingly, Upchurch and Martin informally used C. oxoniensis as the type specimen in their descriptive work, noting that this usage should be formalised by a petition to the International Commission on Zoological Nomenclature (ICZN).

Six years layer, we submitted that petition to the Bulletin of Zoological Nomenclature; a few months after its publication, positive comments from Paul Barrett and Pete Galton followed.

That was in 2009. Five years of silence followed, as the Commission meditated on our five-page petition. (That’s two pages plus front-matter and references). Today, finally, the results are in! The abstract says it all:

The Commission has conserved the usage of the generic name Cetiosaurus Owen, 1841 by designating Cetiosaurus oxoniensis Phillips, 1871 as the type species of Cetiosaurus in place of Cetiosaurus medius Owen, 1842.

So Cetiosaurus finally has a decent type species! Two cheers for the Commission!

I’d always assumed that ratifying the petition would be a no-brainer once the Commission got around to examining it. In fact, their report makes it clear that’s not how it was at all. 16 members voted for the proposal, eight voted against and two abstained. So I guess we were only three switched votes away from having the proposal rejected. Which would frankly have been stupid: every sauropod worker would just have carried right on using C. oxoniensis as though it were the type species anyway.

Why would anyone vote against, you ask? I asked myself the same question. Happily, the decision explains the objections in detail. They nearly all seem to come down to complaints that we didn’t clearly enough explain why C. medius was the previous type species. There’s a reason for that: the truth is that the literature is so vague and contradictory that no-one really knew what the heck the type species was — which is one of the reasons we needed to establish one. (Upchurch and Martin 2002:1053 thought C. brevis was the type; as we investigated this in more detail for the petition, we concluded that the claim of C. medius was stronger. But still very weak.)

But all of that seems like pointless pithering to me. Who cares what the type species was? The point of the petition is to establish what it is, and only one Commission member expressed any reservations about the case we’d made — which is basically that C. oxoniensis is what’s always used in comparisons.

Anyway, dissenting opinions notwithstanding, the genus Cetiosaurus now stands before us having been made an honest woman at long last.

NaishMartill2007-british-dinosaurs-fig3-cetiosaurus

The Rutland cetiosaur, reconstruction by Mark Evans (Naish and Martill 2007: fig 3)

… all of which leaves us with the question of what the Rutland cetiosaur is. It’s been assumed to be Cetiosaurus all along, and that identification has to stand until someone publishes a case to the contrary. But there do seem to be persistent rumours that someone somewhere thinks it’s something different. I wonder if anything will ever come of it?

References

 

How can it be?

IMG_0517

All credit to the Yale Peabody Museum for having the courage to display this historically important object in their public gallery instead of hiding it in a basement. It’s the skull from the original mount of the Brontosaurus (= Apatosaurus) excelsus holotype YPM 1980.

Needless to say, it bears no resemblance at all to the actual skull of Apatosaurus, and the one they now have on the mount is much, much better:

IMG_0500-skull

But how did the YPM people ever arrive at this double-plus-ugly skull above? We see a similar skull in Marsh’s (1891) second attempt at restoring the skeleton of Brontosaurus:

Marsh1891-plateXVI-Apatosaurus-skull

But even this is not as ugly and Just Plain Wrong as the physical model they made. (Marsh’s first restoration of the Brontosaurus skeleton, in 1893, had a much less clear skull.)

So how did the YPM come to make such a monstrosity? What was it based on? Tune in next time for the surprising details!

Bizarrely, we’ve never really featured the  YPM 1980 mount here on SV-POW! — we’ve often shown individual bones, but the mounted skeleton appears only in the background of the much less impressive Morosaurus (= Camarasaurus) lentus mount. We’ll fix that real soon.

 

Last time, we took a very quick look at YPM 1910, a mounted skeleton that is the holotype of Camarasaurus (= “Morosaurus“) lentus, in the dinosaur hall of the Yale Peabody Museum.

Here’s the whole skeleton, in various views. Skip down to the bottom for the science; or just enjoy the derpiness. First, in anterior view:

DSCN8899

Here’s a more informative right anterolateral view. As you can see, this little Camarasaurus is in every sense in the shadow of the the much more impressive Apatosaurus (= “Brontosaurus“) excelsus holotype, YPM 1980: click through for the full image:

DSCN8893-thumbnail

And here’s the corresponding photo from Lull (1930: figure 1) (see below):

Camarasaurus lentus, holotype skeleton, oblique front view (Full 1930: fig. 1)

Camarasaurus lentus, holotype skeleton, oblique front view (Lull 1930: fig. 1)

It’s interesting to see such a familiar mount in such unfamiliar surroundings. Judging by the cabinets in the background, YPM 1910 was mounted in what’s now the dinosaur hall at Yale — i.e. it hasn’t moved since the photo was taken. But back then, Brontosaurus hadn’t been mounted, and Zallinger’s mural hadn’t been painted.

If you thought this animal looked dumb from the front, check out this left anterodorsolateral view, taken from the balcony above the hall. The foreshortening of the neck makes Cam look like a particularly dense puppy.

(Once more, click through for the full version of the photo, including the much more impressive Apatosaurus.)

DSCN8887-thumbnail

Right lateral view, with Zallinger’s justly famous mural in the background. Note the Diplodocus-type double-beamed chevrons in the tail:

IMG_0552

Here’s the justly under-rated posterior view:

IMG_0551

And finally, Lull’s left posterolateral photo — taken from a position that can’t now be replicated, due to the inconveniently located Brontosaurus. (The Archelon in the background, which was previously featured on SV-POW!, has been moved to the end of the hall since Lull’s time.

Camarasaurus lentus, oblique rear view. Lull (1930: fig. 2)

Camarasaurus lentus, oblique rear view. Lull (1930: fig. 2)

How much of this skeleton is real? Happily, not the skull. We can only hope that the real thing wasn’t quite so troubling. But much of the rest of the skeleton is real bone. To quote Lull (1930:1-3):

In the Yale specimen the entire vertebral column is present from the second or third cervical to the tenth caudal with one or two later caudals. Of the limbs and their girdles there are present the left scapula, right coracoid, both humeri, the left radius and ulna, both ilia, the right pubis and left ischium, and both femora, tibiae and fibulae. One cervical rib is present but no thoracic ribs. The disarticulated sacrum lacked one rib from either side.

(How could Lull have been unsure whether the most anterior preserved cervical was the second or third? C2 in sauropods, as in most animals, is radically different from the subsequent cervicals. He does go on to say that only the centrum of the most anterior vertebra is preserved, but the axis has a distinct anterior central articulation.)

Lull is quite ready to criticise the mount, and notes in particular:

The cervical ribs in the Yale mount are not long enough by half, and the thoracic ribs may be somewhat heavy and their length a little short […] both carpus and tarsus are probably incorrect, as the elements in each instance are fewer than shown, there being no more than two at most. There is apparently no justification for the fore and aft extensions of the distal chevrons, as these were not preserved and the Osborn-Mook restoration was followed. […] A probable error lies in too great an allowance for cartilage between the [pelvic] elements, thus making the acetabulum seem rather large.

He also notes a scheme that sadly never came to pass:

[The holotype of Camarasaurus (= “Morosaurus“) robustus], a very perfect specimen, we intend to mount when the great Brontosaurus excelsus type is completed. The three sauropods, ranging in length from 21 to nearly 70 feet, should make a very impressive group.

They would have done! But in the end it fell to the Museum für Naturkunde Berlin to give us the world’s first three-sauropod combo (unless someone knows of an earlier one?)

Finally; the mounted Yale Camarasaurus also crops up in three of the plates of Ostrom and McIntosh (1966). Plate 60 depicts metacarpals I and II in all the cardinal views except for some reason posterior; plate 61 does the same for metacarpals III and IV); and plate 70 shows the right pubis in every aspect but anterior. Here it is:

x

Morosaurus lentus [Now referred to Camarasaurus lentus] Marsh (1889) YPM 1910 (holotype). Right pubis (reversed) in medial (1), posterior (2), lateral (3), proximal (4), and distal (5) views; transverse sections through blade (6) and shaft (7). (Ostrom and McIntosh 1966: plate 70)

Judging by this, it’s a beautifully preserved element with some very distinctive morphology. But we’ve been burned by Marsh’s plates before, and I don’t trust them at all any more — at least, not until I’ve seen the elements for myself. Now I wish I paid more attention to Derpy’s pubes.

And on that line, I’m out.

References

Lull, Richard S. 1930. Skeleton of Camarasaurus lentus recently mounted at Yale. American Journal of Science, 5th series, 19(109):1-5.

Ostrom, John H., and John S. McIntosh. 1966. Marsh’s Dinosaurs: the Collections from Como Bluff. Yale University Press, New Haven, CT. 388 pages including 65 positively scrumptious plates.

Matt’s harsh-but-fair “Derp dah durr” / “Ah hurr hurr hurr” captions on his Giraffatitan skull photos reminded me that there is a sauropod with a much, much stupider head than that of Giraffatitan. Step forward YPM 1910, a mounted skeleton that is the holotype of Camarasaurus (= “Morosaurus“) lentus, in the dinosaur hall of the Yale Peabody Museum. Herp derp derp Full details on this specimen next time! (But a spoiler: the skull isn’t real.)

If your museum doesn't look like this, you should reconsider your existence.

If your museum doesn’t look like this, you should reconsider your existence.

We’re just back from SVPCA 2013 in Edinburgh. The first part of the meeting was held at the Royal Society of Edinburgh, but on Friday we moved to the National Museums Scotland. Which is awesome. And free to the public. The design process for the museum seems to have been, “Okay, let’s get one of, oh, every interesting thing in the world, and put it right here.” We have tons more photos of amazing things from the museum, and maybe we’ll get around to posting them sooner or later, but today I have other things to do.

This pathetic, racially senescent freak is destined for evolution's dustbin.

This pathetic, racially senescent freak is destined for evolution’s dustbin. And he knows it.

Like make fun of Mike. And talk about vomiting dinosaurs.

Dude, this party totally ro-BLAAAUUGGH!!

Dude, this party totally ro-BLAAAUUGGH!!

This groovy stuffed fulmar, Fulmarus glacialis, is shown in the act of puking, which it does to dissuade predators. And probably everyone else. I am reliably informed by Darren that this is unrealistic fulmar vomit, and that the real thing is  more of a thin stream, like the world’s nastiest water gun, which can be directed with considerable accuracy. Note to self: don’t piss off the fulmars.

Vomiting sauropod by Wedel and NichollsLast year cemented “drawing goofy sauropods down at the pub” as a regular SVPCA Thing. So one night I was out with Mike and Darren and paleoartist Bob Nicholls, who is famous around these parts as the creator of the Greatest. Paleoart. Ever. I did a goofy sketch in my notebook illustrating the “defensive vomit” hypothesis, which Brian Engh and I cooked up during this alligator dissection. More on that another time, maybe. Anyway, after bashing out a fairly pathetic sauropod-puking-on-theropod scene, I passed the notebook to Bob and said, “Make this not suck”. Which he did. (Seriously, if you could see my original scrawl, you’d be the one throwing up.)

So now I have an original Bob Nicholls sketch–heck, the world’s first Wedel-Nicholls artist collaboration!–in my notebook, of one of evolution’s most majestic successes responding appropriately to a vulgar, overstudied theropod. Bob drew it right in front of me and I got to drink good beer while I watched him work.

And that, more or less, is why I attend SVPCA.

Giant Irish Mike - cut out

I couldn’t sign off without giving you another version of Giant Irish Mike, with the background cropped out so he can be dropped right into posters, slide shows, and other works of science and art. I really, really hope that he turns up in conference talks and other presentations in the months and years to come. If so, send us a photo documenting his miraculous apparition and we’ll show it to the world.